Most commonly found is pure blue, dark blue, or light blue, although observers find the aqua, purple, or orange variation throughout the ocean. These sea stars may grow up to 30 cm (11.8 in) in diameter, with rounded tips at each of the arms; some individuals may bear lighter or darker spots along each of their arms. Individual specimens are typically firm in texture, possessing the slightly tubular, elongated arms common to most of other members of the family Ophidiasteridae, and usually possessing short, yellowish tube feet. An inhabitant of coral reefs and sea grass beds, this species is relatively common and is typically found in sparse density throughout its range. Blue stars live subtidally, or sometimes intertidally, on fine (sand) or hard substrata and move relatively slowly (mean locomotion rate of 8.1 cm/min).
The genus Linckia, as is true of other species of starfish, is recognized by scientists as being possessed of remarkable regenerative capabilities, and endowed with powers of defensive autotomy against predators.Although not yet documented, L. laevigata may be able to reproduce asexually. L. laevigata is apparently not an exception to this behavior, as many individuals observed in nature are missing arms or, on occasion, in the comet form.
This sea star is fairly popular with marine aquarium hobbyists, where it requires a proper, slow acclimatization before entering the tank system, and an adequate food source similar to that found in its natural habitat. Generally thought of as a detritivore, many sources maintain that this species will indefinitely graze throughout the aquarium for organic films or sedentary, low-growing organisms such as sponges and algae. In the marine aquarium hobby, they have been seen to consume Asterina Starfish, which are commonly introduced into such aquaria on the ubiquitous "live rock" used in such settings.
Lysmata amboinensis is an omnivorous shrimp species known by several common names including the Pacific cleaner shrimp. It is considered a cleaner shrimp as eating parasites and dead tissue from fish makes up a large part of its diet. The species is a natural part of the coral reef ecosystem and is widespread across the tropics typically living at depths of 5–40 metres (16–131 ft).
Lysmata amboinensis can reach a body length of 5–6 centimetres (2.0–2.4 in) and have long white antennae. Mature shrimp are pale in colour with longitudinal bands on their carapace; one central white band with wider scarlet red bands on either side. The shrimp has a long larval stage and unusual sexual maturation, initially being male but becoming a functional hermaphrodite once mature.
Lysmata amboinensis is popular in home and public aquaria where it is commonly referred to as the skunk cleaner shrimp; this is due to its striking colours, peaceful nature, and useful symbiotic cleaning relationship which can also be witnessed in captivity.
Adult shrimp can reach a body length of 5–6 centimetres (2.0–2.4 in) with two pairs of long white antennae. The body and legs are pale amber in colour with longitudinal bands on the carapace: one central white band is flanked by wider scarlet red bands. There are several symmetrical white spots on the red tail. The most anterior antennae fork into two making the shrimp appear to have three pairs of antennae. As with all decapods there are 10 pairs of walking legs, with the most anterior pair having forceps (pincers). Anterior to the walking legs are a pair of white maxillipeds used for feeding. Underneath the thorax behind the walking legs are paired pleopods (swimmerets) used for swimming and brooding eggs.[Lysmata amboinesis have a pair of stalked eyes though experiments suggest they have coarse vision and are colour blind; this low spatial resolution makes it unlikely they can see the striking patterns of other shrimps or tropical fish.
Distribution and habita
Lysmata amboinensis is native to coral reefs across the tropics including the Red Sea and the Indo-Pacific Ocean. They live on tropical coral reefs between depths of 5–40 metres (16–131 ft) and are typically found in caves or reef ledges.
Life cycle
Lysmata amboinesis lay eggs which the adult shrimp keep attached to their pleopods. The early larvae are called nauplii which hatch into more developed larvae called zoeae and go through a free-floating planktonic stage. During this time they feed on other plankton and moult through 14 identified stages growing to approximately 2 centimetres (0.79 in) in length over 5–6 months. At this point the larvae will settle and metamorphose into a more mature form, though not yet the adult state. This is achieved after several moults which occur every 3–8 weeks.
Each shrimp starts out as a male, but after a few moults will become a hermaphrodite allowing them to function as both male and female in interactions with another shrimp; these shrimp have no pure female form. This form of sexual maturation is scientifically described as 'protandric simultaneous hermaphroditism' and is unique to Lysmata shrimp amongst other decapod crustaceans. In one spawning, adult shrimp will lay between 200–500 eggs which are initially attached to the pleopods and are greenish in colour; the eggs swell and lighten in colour before hatching and a few will turn silver on the day of hatching. The eggs hatch around dusk releasing 3–4 millimetres (0.12–0.16 in) long larvae into the free-floating pelagic zone.
Behaviour
Lysmata amboinesis do not live in large groups, more often in pairs, and while omnivorous it is believed they derive much of their nutrition from cleaning parasites and dead tissue from fish. Their mating behaviour has been observed in captivity where it involves little ritual: a pair of fully mature hermaphroditic shrimp will alternate moulting timing, mating occurs shortly following a moult when one shrimp acting as the male will follow the other acting as the female which will brood the fertilised eggs; when the next shrimp moults the roles, and therefore apparent sex, will reverse.
In captivity L. amboinesis have been seen to be socially monogamous showing such aggression that if they are kept in groups of more than 2 individuals one pair will kill the rest. While they are not generally seen in large groups in the wild it is unknown if they are socially monogamous in their natural environment.
Symbiosis
Lysmata amboinesis, like other cleaner shrimp, has a symbiotic relationship with 'client' fish in which both organisms benefit; the shrimp gain a meal from eating parasites living on large fish and the clients benefit from the removal of parasites. Cleaning organisms including shrimp and fish can congregate in one location on a reef in what is called a "cleaning station" which fish will visit when required; cleaning shrimp perform a rocking dance from side to side to signal their services when fish approach. Removal of parasites under captive conditions happens mainly at night though it is unknown whether this is due to shrimp or host fish behaviour.Additionally, cleaning services provided by the shrimp aid wound healing of injured fish supporting the symbiosis hypothesis. L. amboinesis is also commonly found living in caves with their client fish, such as moray eels, providing the shrimp with protection from predators.
Because of the benefits of cleaner shrimp to the fish they clean, Lysmata amboinensis and other species have been suggested as potentially useful to aquaculture.
Common name
Lysmata amboinesis within the marine aquarium trade is commonly referred to as the skunk cleaner shrimp, white-banded cleaner shrimp or scarlet cleaner shrimp due to its appearance, however this last name is also sometimes used for Lysmata debelius.
In the aquarium
Many species of Lysmata, including L. amboinesis, are commonly kept in salt water aquaria; they are safe and beneficial since they will clean both the tank and fish but not harm corals. For these reasons they are often kept in both home and public aquaria for educational purposes, sometimes in 'touch pools' which allow visitors to put their hand in the water so the shrimp will clean their hand.
Despite significant efforts L. amboinesis remains difficult to culture in captivity, though adults will regularly spawn eggs. The cycle from spawning to adult form takes approximately 6 months during which time the vast majority of larvae die. As such most shrimps sold in the aquarium industry are wild-caught, this has led to concern about the negative effects of their removal on natural reefs though there is currently no evidence of an effect.
Lysmata debelius is a species of cleaner shrimp indigenous to the Indo-Pacific. It is popular in the aquarium trade, where it is known as the fire shrimp, blood shrimp or scarlet cleaner shrimp.
Lysmata debelius grows up to 3 centimetres (1.2 in) long, with males and functional hermaphrodites appearing similar. It has a red body, with conspicuous white antennae and white tips to the third to fifth pereiopods. There are white dots on the cephalothorax and legs; white dots on the abdomen distinguish L. splendida from L. debelius.]
Lysmata debelius is one of a group of species in the genus Lysmata that has the role of a cleaner shrimp in reef ecosystems, alongside L. amboinensis, L. grabhami and L. splendida.[5] It is a popular aquarium pet.
Lysmata debelius is a hermaphrodite and therefore any two individuals may mate. The hatching of eggs, moulting, and copulation cycle is identical to that of L. wurdemanni, yielding weekly batches of zoeae from each pair.
